More from reason than from any experience in biology: I don't think macromutation has any statistical chance of being a significant factor in branching of species. The chance of a 'macromutation' both occurring and being successful enough to not only let the spawn survive but even let it thrive as a branched species is vanishingly slim.
Perhaps complex characteristics might evolve as functions with simple gene input, like perhaps the shape of an organ or the structure/color of a fur or even the size of a mammal all expressed by a few genes. This would allow that the species as a whole could adapt quicker to environmental changes. Then the gene mutation would be 'micro' but the resulting change could be 'macro'. Perhaps paedomorphism is like that?
Perhaps complex characteristics might evolve as functions with simple gene input, like perhaps the shape of an organ or the structure/color of a fur or even the size of a mammal all expressed by a few genes. This would allow that the species as a whole could adapt quicker to environmental changes. Then the gene mutation would be 'micro' but the resulting change could be 'macro'. Perhaps paedomorphism is like that?